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Captain Jack Sparrow travels to London to locate a person who has been impersonating him. The imposter has been recruiting a crew to search for the legendary Fountain of Youth. First Sparrow rescues his former first mate, Joshamee Gibbs, who is about to be hanged, though the two are captured. Jack is brought before King George II, who wants him to guide an expedition to the Fountain of Youth before the Spanish find it. Heading the expedition is Jack's old nemesis, Captain Hector Barbossa, now a privateer in service to the British Navy after having lost Jack's ship, the Black Pearl, as well as his leg.

Jack escapes, and is found by his father, Captain Teague, who warns Jack about the Fountain's tests. Jack's imposter is Angelica, his former lover and daughter of the ruthless pirate Blackbeard, who possesses supernatural powers and practices voodoo magic. Jack is forced to join Blackbeard's crew and to lead them to the Fountain. Meanwhile, Gibbs, who memorized, then destroyed Sparrow's map, barters with Barbossa to lead him and his crew to the Fountain.